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There are four
pigeon patterns. Barless (c),
Bar (+), Checker (C), and T-pattern
Checker (CT).
I have the patterns listed from left to right in order of dominance
with barless
being at
the bottom.
Click
on
each
picture
to enlarge them.
The examples above are
given for blue/black color, but patterns can be seen in any of the
base
colors and their diluted and reduced versions. The top dominant is
the T- pattern (gene symbol CT)
which is almost as black (a few light edges or triangles as "check" in
the wing shield area) as spread pigeons. However, T-pattern birds have
regular flights and tail with the near terminal band. By looking at the
terminal tail band, we can determine if the bird is Spread or T-patern
Checker, as spread birds don't have terminal tail band. Also, spread
is not one of the 4 base patterns found in pigeons. Recessive to the
T-pattern is another mutant allele, checker (C), showing several
triangular checks of light gray or "blue" in the blacker wing
shield area. These two types actually intergrade, possibly from modifying
factors
or more likely from intermediate alleles. Nevertheless, one can almost
always distinguish them in pigeons. A third allele recessive to both
T-pattern and check is the wild type bar pattern (+). Finally, the 4th
allele, recessive to the other 3, is the barless (c). Barless is bluish
like the wild type, but it lacks the bars. Barless is only rarely seen
in pigeons.
The gene symbols in order
of decreasing dominance are CT > C > + > c
Like in color, the highest
order of dominance can hide anything below it but nothing above it.
This means that two barred birds cannot produce a T-check or check
offspring. Unlike 3 base colors, there are 4 different traits for
the pattern
and each
gender carry
2 allele for any of the patterns that
are shown above. The pattern therefore is not sex linked because every
pigeon, male or female carry two alleles for patterns. This makes
both sexes homozygous or heterozygous for any of the patterns. A
hen can
be heterozygous
bar carrying barless or homozygous T-pattern checker (both alleles
for pattern is T-pattern checker). But she cannot be heterozygous bar
carrying
checker! During
mating, each gender donate one of the alleles for the pattern that they
carry and the
offspring will then show the dominant pattern
it received from both of its parents. If the offspring receives same
allele patterns (both bar, or both checker) then the pattern would
be homozygous.
It is very important
to note that the pattern is located in different locus (different
forms of a gene at a particular physical location on
a chromosome) and therefore inherited independently of color. The pattern
does not occur only on blue but in every other color except recessive
red, spread black, and white. Understanding the pattern and their
inheritance is very
important to breeders interested in color, specially to the breeders
of Toy Stencils, Oriental Frills, etc. According to Wendell Levi,
in his book The Pigeon, "when breeds or varietties of different
color are crossed, blue color frequently appears. This accurence,
called
"reversion", was one of the
Darwin's chief reasons for believing that all domestic pigeons are
descendent
from the blue rock pigeon (C. livia)" (Levi, p. 309).
People often ask me how
they can get barless birds. Barless is a mutation and a unit that
you have to have in order to produce more offspring.
In addition, being at the bottom of the dominance it takes couple of
generations of inbreeding to get it back. At the moment I have a pair
of barless rollers but I am not very happy with their performance.
So, my
goal is to breed them with better performers and then breed the offspring
back to the barless parent to keep the barless pattern.
Here
is how to breed barless from
one parent:
Let's
say I have a barless hen. She must be homozygous barless in order
to
show a barless pattern, as
barless is expressed only in homozygous stage and her genetic symbol
would be represented as c//c. Now, if I crossed her with a homozygous
bar cock, both alleles for pattern is bar and represented as +//+.
All of the offspring from that mating will be heterozygous bars carrying
barless. Now, if I cross the male offspring from that mating, back
to his barless mother I would get: 50% barless and 50%
heterozygous bar carrying barless. The punnet square shown
on the left is the demonstration of this mating. By keep
inbreeding them, I can keep the barless pattern and add performance
to them.
Unfortunately,
the color and the pattern are easier to put in to a breed than the
performance, because performance in pigeons is also a recessive trait
and very hard to put in and maintain. Think of performance
as a talent
that not every pigeon is blessed with.
Barless is a fairly rare
mutant in most domestic pigeon breeds and often confused with other
modifiers. I have seen a lot
of people calling magpies,
homozygous indigo (andalusions), recessive reds even white-selves (I
actually heard that one!) as barless. I have also seen people calling
a T-pattern
checker as spread. It is actually very easy to separate a spread from
a T-pattern by just looking at their tail band. T-pattern Checker,
Checker, Bar and Barless birds will have tail bands
and you can actually see a white line on the edges of end tails;
unless the birs is an ash red. Ash red is a color where the tail
bar is not brick red that we would see in the bars of the pigeon.
Instead, we find that it has been washed out to an ashy-gray color
just like the color
that we found on the edges of the wing's flight feathers.
Here
is how to breed bar pattern from two chequer parents:
I
once received an email from someone wondering how in the world he
got a bar pattern from
two chequer parents. The answer is very simple. In order for two
checkers to produce a bar pattern, both parents have to be heterozygous
chequers carrying bars. Both parents must be carrying a bar pattern
in order to produce a less dominant bar pattern from two
chequer parents. From this mating there is 25% chance or 1 out of
4 babies will be a bar pattern.
The
punnet
square
on
the
right
demonstrates
how this is possible. 25% of this mating will be Homozygous Chequer
(both alleles for patterns are chequers) as it is
shown on top left sequare. In addition, 50% of the offspring from
this
mating
will
be Heterozygous Chequers carrying Bar just like the parents, as it
is shown on top right and bottom left squares.
Other
modifiers and markings:
A
pigeon may not show its true pattern because other
modifiers prevent it like white-self or
any other spread color i.e. black, recessive red or
brown. Andalusion, grizzle, almond, toy stencil, lace, magpie or white
side birds will also be preventing for us the see the pattern of the
bird, but understand that the pattern is there (under the modifier).
Notice that other than the 4 patterns shown above, the other modifiers
like
spread
black or
white-self is NOT considered to be pattern or color but as a modifiers
and they are located in a different chromosome than the pattern and
the
color.
If a bird has a spread gene in it, it will basically cover (hide) the
pattern of the pigeon and will make the pattern not visible. But, regardless
of which modifier is expressed, a pigeon still carries two additional
alleles for the pattern under a spread whether it is homozygous or heterozygous.
In other words, every spread bird we see, still carries additional two
out of four patterns (barless, bar, checker, and T-pattern checker)
in
them
which
are simply covered (modified) by the spread modifier.
The following pictures
show most of the modifiers that prevent us seing the 4 base patterns.
Click on each picture to enlarge them. |
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